In the previous blog, I discussed very briefly how spending some time in the company of the same individual or small group of animals on separate occasions has allowed me to explore their lives more intimately and meaningfully through photography. This is one strategy that I relish whenever I stumble across such an infrequent and special instance. I indicated too that spending time repeatedly in the same small locality on separate trips has allowed me to maximise the odds of capturing the sense of place much more intensely. Repeated visits have not only allowed me to broaden my understanding, appreciation and feel for a locale; recurring outings to the same area also spur me on to work harder at capturing all features of a place photographically.
The Nature of Nature - African wildlife and environmental images, stock photography and fine art prints.
Showing posts with label grasslands. Show all posts
Showing posts with label grasslands. Show all posts
Sunday, 6 December 2020
ON PHOTOGRAPHY: Maximising the Odds of Photographic Expression – 2. A Farm Called Palmietpan
In the previous blog, I discussed very briefly how spending some time in the company of the same individual or small group of animals on separate occasions has allowed me to explore their lives more intimately and meaningfully through photography. This is one strategy that I relish whenever I stumble across such an infrequent and special instance. I indicated too that spending time repeatedly in the same small locality on separate trips has allowed me to maximise the odds of capturing the sense of place much more intensely. Repeated visits have not only allowed me to broaden my understanding, appreciation and feel for a locale; recurring outings to the same area also spur me on to work harder at capturing all features of a place photographically.
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Tuesday, 14 July 2020
A GRASS MENAGERIE: II - The Inhabitants of the Grasslands
In
essence, the smaller denizens of the grasslands (including the myriad of
insects inhabiting this biome) live in a forest-like environment – not under a
canopy of leaves and between tree trunks, but rather between the culms and the
inflorescences of the taller grasses. Their environment is dense and provides
for ample opportunities either to shelter or to escape from predators.
Moreover, these smaller organisms, experience subtle stratification of the
environmental factors (such as light intensity, temperature and humidity) from
ground level upwards to the top layer of the grasses. These littlies can move
up and down in their environment to modify, albeit very slightly, the influence
on their tiny bodies of a few environmental factors. For example, any small
motile animals living in the grasslands can escape the heat of a summer’s day by
moving down to the shaded ground level; they can escape desiccating winds, again
by sheltering at the bases of the grasses. Despite these advantages, the
differences between day and night are significant and can prove to be
challenging to the small inhabitants; at night the grasslands become much
colder, as a lot of heat is re-radiated to outer space, while far less heat is
trapped in the sparse biomass of the vegetation. Of course, for the larger
animals the grasslands are essentially two-dimensional environments, with the
subtle changes in environmental conditions proving insignificant (apart from
changes from day to night).
For
many animal species, the grasslands become uninhabitable during the winter
seasons. In general, two responses have evolved to this challenge. Those motile
organisms that can not cope with the prolonged dry, cold times will migrate to
areas that are more amenable in environmental conditions. The larger herbivores
often partake in long-distance migrations, following the patchy and
intermittent rainfall. Many bird species are trans-equatorial migrants, using
long-distance flight to inhabit grasslands only during the wetter summer months
in both hemispheres.
Many
of the smaller organisms that are not able to escape from the rigours of the
winter season by migration over large distances enter a period of dormancy,
often for significant stretches of time. In regions of the planet with warmer
climates, many insects living exclusively in grasslands aestivate during the most
arid months. Populations of active insects will fluctuate in number seasonally;
moreover, the occurrence of large, stable populations of insects in local areas
of grasslands tends to be less predictable than in other biomes. A number of
insect taxa do cope well with the environmental conditions in grasslands. These
include orders of insects such as the Orthoptera (including grasshoppers and
locusts) and the Hymenoptera (including ants). The very successful insects
consume the leaves of grasses themselves, or they harvest, store and consume
the grass seeds.
For
the much larger vertebrate herbivores that inhabit the grasslands, food is
present in abundance, provided the animals can graze the grasses themselves,
feed on the grass seeds or select the herbs nestling deeper down between the
grasses. Although the biodiversity of grassland plants is generally very high,
the vegetation types available for herbivores is limited to grasses and
smaller, usually non-woody flowering plants that grow here. Consequently, the
diversity of animal life that is supported by grasslands is impoverished (in
general) when compared to other biomes like the woodlands and forests, at least
in terms of the number of species that inhabit the grasslands. This is not
necessarily the case in terms of the number of individuals that live in the
grasslands, since adaptation to this periodic environment has occurred in many
groups of organisms (such as the grasses themselves, and the flock-forming
seed-eating birds and the herd-forming grazing antelopes).
Amongst
the numerous antelope species living in grasslands and open savannas, an
interesting correlation exists between the size of individuals of a species and
the social behaviour that has evolved in that species. The smaller browsers and
grazers require vegetable food sources of high nutritional quality. Their
bodies have a small volume compared to their surface area. To keep fuelling the
high rate of cellular respiration required to maintain a constant and high core
body temperature, these small antelope must be highly selective in the food
that they consume. Therefore, small antelope species are solitary or live in
small family groups.
In
the much bigger herbivores, the quality of the food is not that vital. Inherent
in the large size is a decreased ratio between the surface area of the organism
and its volume that allows for a slower basal metabolic rate than that found in
smaller species. With a small surface area compared to volume, large organisms
are not faced with excessive heat loss through their skin. At the same time,
the larger volume allows for the digestion of the food to proceed at a slower
pace. Large antelope can graze grasses and allow the digestion to take place
over time; hence the evolution of multi-chambered stomachs and the behaviour of
chewing the cud in the antelope species. The alternative strategy, seen in
horses and zebras, for example, involves the evolution of a large caecum; in essence,
this is a spacious storage vat in the hindgut in which vegetation can be
decomposed by bacterial action. Whatever the anatomy of these larger
herbivores, they can survive easily in vast herds since the food they consume
is more easily available, albeit of low nutritional value.
Add
to this another benefit: the relatively low height of the grassland vegetation
allows for vigilance against predators for the larger animal species. Living in
a large group means more eyes and ears, as well as a smaller chance that a
particular attack by predators will result in the death of any one of the
members of the group. (Of course, one of us will die, but the chance of it
being I is lessened in a large group.) The ability to live in larger herds has
necessitated some form of herd co-operation and control that is complex enough
to promote social cohesion, especially during courtship and mating. So vast
herds of large grazers occupy the grassland environment, tracking the
intermittent rainfall in spectacular long-distance migrations. Their predators,
too, follow the migrations, but to a lesser extent – territoriality between
carnivores well-equipped with the weapons of their trade can be intense and will
limit the movement of groups of predators.
Another
feature vital for the survival of mammalian herbivores in grasslands is the
production of precocial young, that is, babies that are fully functional very
soon after birth. The gestation periods of grassland herbivores is lengthened,
in general, and the newborn is able to be on its feet and run with the herd
within minutes after birth. In the smaller herbivores, where no herd behaviour
is found, the young are precocial nevertheless, although they will often hide
in denser vegetation or denser patches of grassland near where the mother is
feeding.
The
lack of tall vegetation in which predators can conceal themselves has
necessitated the evolution of larger carnivores that can run down their prey,
rather than relying on concealment and a pounce at close quarters. Three
strategies in hunting prey among the larger carnivores prevail in grasslands:
sheer speed of pursuit, endurance of long-distance chasers or the use of
elaborate hunting strategies in larger social units of predators. Predators
that rely only on speed still live either as solitary individuals or in small
family groups, usually a female and her offspring only. This is often not the
case with the two other groups – the long-distance chasers and the strategic
hunters both require the co-operation of several members of the group in order
to kill prey. This interdependence of members of a group for effective hunting
goes hand in hand with more complex and elaborate social behaviour. Maintaining
co-operation in a pack or pride requires social interactions that promote group
cohesion, such as appeasement behaviours and social ranking of individuals
within the group. Invariably, a hierarchical social system develops in these
predatory species.
For
the rodents, the omnivores and the smaller carnivores of the grasslands,
shelter from the harsh environment, concealment from predators, or safety
during reproduction is often found in burrows. The grassland environment, with
its relatively deep soils and the top layer of the soil knitted together by the
adventitious root systems of the grasses allows deep tunnel systems to be
excavated by many animals that have the strength and the equipment (such as
large claws) to do so. Subsequent occupation of the burrows by a host of other
species – comprising individuals that themselves can not dig effectively –
means that these tunnel systems often stay in use for long periods.
Photographing wildlife in grasslands is challenging. The larger species
(mammals and birds) will flee readily and they can not be approached without
difficulty. The unpredictability of the occurrence in any specific area of
larger animal species can be taxing too. For me, portraying the grasslands has
remained a test that I accept with glee. To display the magnificence of this
biome and its menagerie of inhabitants will remain a special privilege always –
particularly so since I too am an original inhabitant of the Highveld’s grasslands.
Friday, 10 July 2020
A GRASS MENAGERIE: I - The Environment of the Grasslands
The
world’s grasslands flourish in the middle latitudes of our planet, quite far
north and south of the equator; hence, this biome is subjected to pronounced
seasonal changes in environmental factors. Grasslands also tend to thrive in
the drier interior of continents some distance from the oceans.
Grasslands
generally receive moderate, often erratic rainfall, usually in late spring to
early summer; the short wet season is followed by prolonged dry periods during
which very little precipitation falls. Temperatures in grasslands can fluctuate
annually from hot daytimes to very cold winters with significant bouts of frost
(at least). Moreover, most grasslands occur on deep soils (often sandy or friable)
that do not retain soil moisture for long.
The
grasslands survive pronounced seasonal changes in a combination of environmental
factors: sufficient warmth and sufficiently long day-lengths for growth, moist
upper layers of soil, and frost-free nights in the summer months, with cold and
frosty nights, dry soils, and short day-lengths in winter. This combination of
factors gives rise to an obvious growing season for the vegetation in late
spring and summer, followed by a lengthy dormant season.
In
southern Africa, the grasslands are located chiefly in South Africa (on the Highveld (an elevated central plateau)
and the inland areas of KwaZulu-Natal and the Eastern Cape) with much smaller,
localised regions found particularly in the eastern highlands of Zimbabwe. In
general, the topography is flat or undulating, but it includes the escarpment areas
of the Highveld and the highlands of
Zimbabwe too. The altitude of southern Africa’s grasslands varies from near sea
level to close to 3000 metres in elevation.
A
copious number of grass species dominates the grasslands extensively. Between
the grasses also grow many non-woody flowering plants and many geophytes
(bulb-producing plants); regularly woody shrublets can also survive the
environmental and climatic conditions experienced in grasslands. Since the
height of the vegetation is very low, plants in grasslands experience high
light intensities (at least during the summer months); no true shade-plants
survive here.
Taller
shrubs and trees are generally absent from grasslands. There are several
reasons for this absence. The deep and often well-drained soils of grasslands
frequently become too dry at deeper levels for tree roots to be able to absorb
sufficient moisture. In general, the leaves and smaller twigs of trees and
shrubs are also not frost resistant; therefore, taller woody vegetation can not
survive the lengthy cold, dry winter seasons. During the winters, fire too
becomes a significant ecological factor in grasslands, removing dense old growth
from the biome, making nutrients available again for the rapid growth of new
grasses during the following growing season. Fire also prevents the
encroachment of shrubs and trees into grassland areas.
In
contrast to most woody vegetation, grasses can survive the challenging
conditions of the dry seasons during which grasses become dormant. In winter, the
grasses usually die back; that is, the above-ground culms dry out completely
while the below-ground root systems survive the harshest of parched cycles. Alternatively,
many grasses are annual plants rather than perennials – the adult lives for one
season only, during which time the plant germinates from seed, grows rapidly,
flowers and sets seed itself, only to die when the dry season commences. As
long as the upper soil layer does not become too hot during a fire, the seeds
of the previous summer can survive there and the grasslands can persist even through
regular cycles of wildfires. The
smaller non-woody herbs, flowering plants and the shrubs of the grasslands
follow similar strategies.
Grasses
form a special subset of the vegetable world in many ways. Their roots do not
penetrate deep into the soil; rather, the many thin, fibrous roots spread out
just under the soil surface, knitting together the top layer of soil. The
decomposition of the dead leaves, culms and roots of grasses releases nutrients
into the topmost, enmeshed and very fertile soil layer. The web of fibrous
roots also captures much soil water before it percolates into the deeper strata
following good rains. Sacrificing a long life for rapid growth, grasses with
their strap-like leaves and thin culms have done away with thick trunks and
large leaves. The strategy employed is one of grow, reproduce and set seed as
rapidly as possible. In grasslands, plants that do not produce seeds are very
rare.
Although
they are flowering plants, grasses have done away with large, showy flowers.
The grasslands, in general, are often too dry for much of the year to support a
permanent, large insect population. There exist vast numbers of insects in
grasslands, but their appearance in a local area may be unpredictable.
Therefore, attracting a pollinator at the correct time – when the flowers have
grown and matured, and are ready for pollination and fertilization – is a
challenge for any flowering plant in a grassland environment. Instead, grasses
have reverted to wind pollination and the flowers have become minute, but
bright, splashes of colour in the inflorescences of grasses. Since most grasses
grow to a more or less uniform height and the grass stalks are thin, wind can
move over and through grasslands relatively unimpeded, except at ground level.
Thus wind pollination is an effective strategy for plant reproduction in this
environment.
In
contrast to all other plants, the grasses possess a unique feature. In plants
that are not grasses, cell division for growth of the organism takes place only
in two regions of the plant: at the tips of the roots and twigs (where
elongation of all plants takes place), and in woody plants just below the outer
covering inside of the bark of larger shrubs and trees (so that an increase in
the diameter of roots, trunks and branches can take place). In grasses,
however, patches of undifferentiated tissues (the intercalary meristems) occur
and cell division can take place at any regular point along the plant. As a
consequence, grasses do not stop growing when their leaves and culms are grazed
(that is, when the tips of the plants are removed); rather, growth of new plant
material will take place from the intercalary meristems. In fact, grasses tend
to thrive better if they are grazed to a certain extent. The saliva of the
antelopes that graze the grasslands of Africa and elsewhere also stimulates
grass growth, an interesting example of co-evolution between a predator and its
prey in these two groups that evolved on our planet at the same time.
When
people look at grasslands, they may conclude that the biodiversity of this
biome is quite low. Grasslands look uniform – there seem to be only grasses
present, and these tend to grow to the same length (more or less) in any
particular area. However, in South Africa, this biome harbours a tremendously high
biodiversity (second only to the fynbos biome of the southern Cape). The
grasslands are often home to rare plants, particularly so in the higher-lying areas
along the escarpment. These scarce species are often endangered; they comprise
mainly endemic geophytes (bulb-producing plants) and dicotyledonous herbaceous
plants. However, very few species of grasses are rare or endangered, almost
certainly as a result of wind pollination and the production of small seeds.
All
my life I have been a resident of the Highveld
of South Africa. My earliest recollections of wilderness are intertwined
intimately with the grasslands. As a very young photographer, I often lamented
the fact that my family had not lived in a bushveld or coastal area –
environments that seemed so much more photogenic in themselves and that
appeared to be jam-packed with opportunities of capturing images of wildlife. However,
the more I explored, the closer I looked at the grasslands surrounding my home,
the more grew my appreciation of the spectacular expanses of the grasslands.
Nowadays, I relish the challenge of portraying the grasslands in all their
splendour; I cherish every opportunity of arresting, in an image, the fabulous grass
menagerie.
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